1. School of Human Development and Health, Faculty of Medicine, Southampton General Hospital University of Southampton Tremona Road Southampton SO16 6YD UK
2. Department of Plant Sciences Rothamsted Research West Common Harpenden AL5 2JQ UK
3. Centre for Biological Sciences, Faculty of Natural and Environmental Sciences University of Southampton University Road Southampton SO17 1BJ UK
4. NIHR Southampton Biomedical Research Centre University Hospital Southampton NHS Foundation Trust and University of Southampton Tremona Road Southampton SO16 6YD UK