1. Department of Earth, Ocean and Atmospheric SciencesUniversity of British ColumbiaVancouver British Columbia Canada
2. Now at Department of Marine Chemistry and GeochemistryWoods Hole Oceanographic InstitutionWoods Hole Massachusetts USA
3. Fisheries and Oceans CanadaInstitute of Ocean SciencesSidney British Columbia Canada
4. Department of BotanyUniversity of British ColumbiaVancouver British Columbia Canada
5. Center for Advanced Science and TechnologyTokyo University of Marine Science and TechnologyTokyo Japan