1. Department of Plant Science and Environmental Ecology; Faculty of Resource Science and Technology; Universiti Malaysia Sarawak 94300 Samarahan, Sarawak Malaysia
2. School of Biological Sciences; Universiti Sains Malaysia; 11800 Penang Malaysia
3. Forest Herbarium (BKF); The Office of Forest and Plant Conservation Research; National Park, Wildlife and Plant Conservation Department; 61 Phahonyothin Road, Chatuchak Bangkok 10900 Thailand
4. Institute of Biodiversity and Environmental Conservation; Universiti Malaysia Sarawak; 94300 Samarahan, Sarawak Malaysia