1. Department of Biology; Queen's University; Kingston Ontario Canada
2. Department of Biology; Tibet University; Lhasa China
3. Institute of Biodiversity Science, School of Life Sciences; Fudan University; Shanghai China
4. Key Laboratory of Yangtze River Water Environment, Ministry of Education, College of Environmental Science and Engineering; Tongji University; Shanghai China
5. Department of Environmental Science, Saint Mary's University; Halifax Nova Scotia Canada