1. United Graduate School of Agricultural Science, Kagoshima University, Kagoshima, Japan
2. Faculty of Agriculture, Department of Bioscience and Biotechnology, University of the Ryukyus, Okinawa, Japan
3. Okinawa Prefectural Agricultural Research Center, Okinawa, Japan
4. Faculty of Agriculture, Department of Biochemical Science and Technology, Kagoshima University, Kagoshima, Japan