1. Yunnan Key Laboratory for Palaeobiology Yunnan University 650500 Kunming China
2. Institute of Palaeontology Yunnan University 650500 Kunming China
3. MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment Yunnan University 650500 Kunming China
4. School of Geography, Geology and the Environment University of Leicester University Road Leicester LE1 7RH UK
5. Chengjiang Fossil Museum of the Management Committee of the Chengjiang World Heritage Fossil Site Chengjiang China
6. Centre de recherche en Paléontologie—Paris (CR2P), CNRS, MNHN Sorbonne Université, Muséum National d'Histoire Naturelle CP38, 57 rue Cuvier F‐75005 Paris France
7. Southwest United Graduate School 650000 Kunming China
8. Natural History Museum London SW7 5BD UK