1. Faculty of Life and Environmental Sciences University of Tsukuba 1‐1‐1 Tennodai Tsukuba Ibaraki 305‐8571 Japan
2. Makino Herbarium Tokyo Metropolitan University 1‐1 Minami‐osawa, Hachioji Tokyo 192‐0397 Japan
3. Faculty of Symbiotic Systems Science Fukushima University 1 Kanayagawa, Fukushima Fukushima 960‐1296 Japan
4. National Research Institute of Fisheries Science, Japan Fisheries Research and Education Agency 2‐12‐4 Fukuura, Kanazawa Yokohama Kanagawa 236‐8648 Japan
5. Sugadaira Research Station, Mountain Science Center University of Tsukuba 1278‐294 Sugadaira‐kogen Ueda 386‐2204 Japan