1. Key Laboratory for Molecular Biology and Biopharmaceutics, School of Life Science and Technology Mianyang Normal University Mianyang Sichuan China
2. Ecological Security and Protection Key Laboratory of Sichuan Province Mianyang Normal University Mianyang Sichuan China
3. Centre for Integrative Ecology, School of Life and Environmental Sciences Deakin University Geelong Vic Australia
4. Ecological Genetics Research Unit, Organismal and Evolutionary Biology Research Programme, Faculty Biological & Environmental Sciences University of Helsinki Helsinki Finland