1. Graduate School of Fisheries and Environmental Sciences; Nagasaki University; 1-14 Bunkyo-machi Nagasaki 852-8521 Japan
2. Institute for East China Sea Research, Organization for Marine Science and Technology; Nagasaki University; 1551-7 Taira-machi Nagasaki 851-2213 Japan
3. College of Oceanography; Hohai University; 1 Xikang Road, Gulou District, Najing China