1. CEFE, CNRS, Univ Montpellier, Univ Paul Valéry Montpellier 3, EPHEIRDMontpellier France
2. Institut des Sciences de l'Évolution, Université de Montpellier, CNRS, IRDEPHEMontpellier France
3. Institute of Evolutionary Biology, School of Biological SciencesUniversity of Edinburgh Edinburgh EH9 3JT United Kingdom
4. INRAUR 0629 URFMF‐84914 Avignon France
5. Univ. Bordeaux, INRAE, BIOGECOBordeaux France
6. Université Bourgogne Franche‐ComtéUMR 6249 Chrono‐environnement16 route de Gray, F‐25030 Besançon Cedex France
7. CNRS, Biodiversity Research CenterUniversity of British ColumbiaVancouver Canada