1. Key Laboratory of Vertebrate Evolution & Human Origins of Chinese Academy of Sciences Institute of Vertebrate Paleontology & Paleoanthropology Chinese Academy of Sciences Beijing 100044 China
2. CAS Center for Excellence in Life & Paleoenvironment Beijing 100044 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Division of Paleontology American Museum of Natural History New York 10024 USA
5. CAS Center for Excellence in Tibetan Plateau Earth Sciences Beijing 100101 China
6. Tianjin Natural History Museum Tianjin 300201 China