1. University of Florida Herbarium (FLAS) Florida Museum of Natural History Gainesville Florida 32611 USA
2. Department of Research, Conservation and Collections Desert Botanical Garden Phoenix Arizona 85008 USA
3. College of Liberal Arts and Sciences Arizona State University Tempe Arizona 85287 USA
4. Desert Solitaire Botany and Ecological Restoration San Diego California 92103 USA
5. School of Biological Sciences Southern Illinois University Carbondale Illinois 62901 USA