1. Graduate School of Integrated Sciences for Life, Hiroshima University, Hiroshima 739-8526, Japan
2. Amphibian Research Center, Hiroshima University, Hiroshima 739-8526, Japan
3. Faculty of Life and Environmental Sciences, Shimane University, Shimane 690-8504, Japan
4. Division of Developmental Biology, School of Life Sciences, Faculty of Medicine, Tottori University, Tottori 683-8503, Japan