1. College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510642, China; and Guangdong Engineering Research Center for Grassland Science, South China Agricultural University, Guangzhou 510642, China
2. College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510642, China
3. Department of Horticulture, University of Arkansas, 316 Plant Science Building, Fayetteville, AR 72701