1. College of Forestry, Nanjing Forestry University, Nanjing 210037, China, and Jiangsu Qinghao Landscape Horticulture Ltd. Co., Nanjing 211225, China
2. Wuping Golden Branch Rare Flowers Ltd. Co., Longyan 364399, China
3. Department of Horticulture, University of Georgia, Athens, GA 30602
4. College of Forestry, Nanjing Forestry University, Nanjing 210037, China
5. College of Forestry, Nanjing Forestry University, Nanjing 210037, China; Jiangsu Qinghao Landscape Horticulture Ltd. Co., Nanjing 211225, China; and National Holly Engineering Center, Hangzhou 311300, China
6. College of Horticulture and Gardening, Yangtze University, Jingzhou 434025, China