1. Department of Agriculture, Central University of Technology, Free State, Private Bag 6 x20539, Free State, Bloemfontein, 9300, South Africa
2. The Agricultural Research Council, Grain Crops Institute, Private Bag X1251, Potchefstroom, 2520, South Africa
3. Department of Agriculture and Animal Health, College of Agriculture and Environmental Sciences, University of South Africa, Private Bag X6, Florida 1710, South Africa
4. Earth and Environmental Sciences, School of Agricultural, University of Kwa-Zulu Natal, Private Bag X01, Scottsville, Pietermaritzburg 3209, South Africa