1. Laboratory of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, PR China
2. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, 266071, PR China
3. Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology (Qingdao), 266237, PR China
4. University of Chinese Academy of Sciences, Beijing 100049, PR China
5. Key Laboratory of Marine Sedimentology and Environmental Geology, First Institute of Oceanography, MNR, Qingdao 266061, PR China