1. Division of Vertebrate Zoology, Department of Mammalogy, American Museum of Natural History
2. Instituto Argentino de Nivología, Glaciología, y Ciencias Ambientales, CCT–CONICET, Mendoza, Argentina
3. Division of Vertebrate Zoology, Department of Mammalogy, American Museum of Natural History; Division of Evolutionary Ecology, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland