1. Molecular Ecology and Evolution Group, Biosciences College of Life and Environmental Sciences, University of Exeter Exeter EX4 4QD, UK & Field Conservation and Research Department, Whitley Wildlife Conservation Trust Paignton Zoo, Totnes Road, Paignton TQ4 7EU, UK
2. Southern Tanzania Elephant Program Box 2494, Iringa, Tanzania
3. Tropical Biodiversity Section, MUSE-Science Museum Corso del Lavoro e della Scienza 3, 38123 Trento, Italy & Udzungwa Ecological Monitoring Centre c/o Udzungwa Mountains National Park, P.O. Box 99, Mang'ula, Tanzania
4. Department for Food and Resource Economy, Faculty of Science University of Copenhagen, Copenhagen, Denmark
5. Whitley Wildlife Conservation Trust, Paignton Zoo Totnes Road, Paignton TQ4 7EU, UK
6. Molecular Ecology and Evolution Group, Biosciences College of Life and Environmental Sciences, University of Exeter Exeter EX4 4QD, UK