1. Cancer Genomics and Immunology Group, Wellcome Centre for Human Genetics University of Oxford Oxford UK
2. Oxford NIHR Comprehensive Biomedical Research Centre, Oxford University Hospitals NHS Foundation Trust Oxford UK
3. Department of Pathology Leiden University Medical Center Leiden The Netherlands
4. Gastrointestinal Stem Cell Biology Laboratory, Wellcome Centre for Human Genetics University of Oxford Oxford UK
5. Department of Otorhinolaryngology, Head and Neck Surgery LMU Klinikum Munich Germany
6. German Cancer Consortium (DKTK), Partner Site Munich Germany
7. Department of Molecular and Cellular Biology Baylor College of Medicine Houston TX USA
8. Reproductive and Developmental Biology Laboratory National Institute of Environmental Health Sciences Research Triangle Park NC USA
9. Department of Life Sciences, College of Health, Medicine and Life Sciences Brunel University London Uxbridge UK
10. Institute of Genetics and Cancer The University of Edinburgh Edinburgh UK
11. Oxford Cancer Centre, Churchill Hospital Oxford University Hospitals Foundation NHS Trust Oxford UK