1. Laboratory of Infection and Prevention Institute for Virus Research Kyoto University Sakyo‐ku Kyoto Japan
2. CREST JST Sakyo‐ku Kyoto Japan
3. Laboratory of Host Defense WPI Immunology Frontier Research Center (IFReC) Osaka University Suita, Osaka Japan
4. Research Institute for Microbial Diseases Osaka University Suita Osaka Japan
5. Laboratory of Allergic Diseases Institute for Advanced Medical Sciences Hyogo College of Medicine Suita Osaka Japan
6. Laboratory of Systems Immunology WPI Immunology Frontier Research Center (IFReC) Nishinomiya Hyogo Japan
7. IBMC UPR 9022 CNRS Strasbourg Cedex France