1. Biological Laboratory, College of Bioresource Sciences, Nihon University
2. Atmosphere and Ocean Research Institute, The University of Tokyo
3. Atmosphere and Ocean Research Institute, The University of Tokyo:Section of Ecology, Department of Biology, University of Turku
4. Kagoshima Prefectural Fisheries Technology and Development Center
5. Laboratory of Marine Environmental Microbiology, Division of Applied Biosciences, Graduate School of Agriculture, Kyoto University