1. Marine Bioresource Science, Division of Marine Bioresource and Environmental Science, Graduate School of Fisheries Science, Hokkaido University:Ecologically Related Species Section, Tropical Tuna Resources Division, National Research Institute of Far Seas Fisheries
2. COE for Neo-Science of Natural History, Graduate School of Fisheries Sciences, Hokkaido University
3. Marine Bioresource Science, Division of Marine Bioresource and Environmental Science, Graduate School of Fisheries Science, Hokkaido University