1. Jiangsu Key Laboratory for Horticultural Crop Genetic Improvement, Institute of Horticulture, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China
2. Department of Plant Sciences, University of Tennessee, Knoxville, TN 37996, USA
3. Department of Entomology and Plant Pathology, University of Tennessee, Knoxville, TN 37996, USA