1. Department of Developmental and Regenerative Biology, Medical Research Institute, Tokyo Medical and Dental University, Tokyo 113-8510, Japan
2. Department of Molecular and Systems Neurobiology, Graduate School of Medicine, University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan
3. Global COE Program “Comprehensive Center of Education and Research for Chemical Biology of the Diseases”, University of Tokyo, Tokyo 113-0033, Japan