1. Laboratory of Clinical Veterinary Medicine for Large Animal, Division of Veterinary Medicine, School of Veterinary Medicine, Kitasato University, Towada 034-8628, Japan
2. Department of Marine Bioscience, Atmosphere and Ocean Research Institute, The University of Tokyo, Kashiwa 277-8564, Japan
3. Hanwoo Research Institute, National Institute of Animal Science of Korea, Pyeongchang-gun, Gangwon-do 25340, Republic of Korea
4. Laboratory of Theriogenology, Department of Veterinary Clinical Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo 060-0818, Japan
5. Laboratory of Animal Reproduction, Department of Animal Science, School of Veterinary Medicine, Kitasato University, Towada 034-8628, Japan