1. Institute of Ageing and Chronic Disease, William Henry Duncan Building, 6 West Derby Street, Liverpool L7 8TX, UK
2. Centre for Genomic Research, Institute of Integrative Biology, Biosciences Building, Crown Street, University of Liverpool, Liverpool L69 7ZB, UK
3. Department of Orthopaedic Surgery, Maastricht University Medical Centre, 6202 AZ Maastricht, Netherlands