1. Department of Applied Ecology, Saint-Petersburg State University, 16 Line of Vasilyevsky Island, Saint Petersburg 199178, Russia
2. Institute of Biology KomiSC UrB RAS, 28 Kommunisticheskaya Street, Syktyvkar, Komi Republic 167982, Russia
3. Arctic and Antarctic Research Institute, 38 Beringa Street, Saint-Petersburg 199397, Russia