1. The Arnold and Mabel Beckman Macular Research Center at the Doheny Eye Institute, University of Southern California, Los Angeles, CA 90033, USA
2. Department of Pathology, Keck School of Medicine, University of Southern California, Los Angeles, CA 90033, USA
3. Department of Ophthalmology, Keck School of Medicine, University of Southern California, Los Angeles, CA 90033, USA
4. Department of Pharmaceutical Sciences, School of Pharmacy, University of Southern California, Los Angeles, CA 90033, USA