1. Laboratory of Biochemistry of Microorganisms, Department of Biochemistry, State University of Maringá, Avenue Colombo 5790, 87015-900 Maringá, PR, Brazil
2. Federal Institute of Paraná, Campus Jacarezinho, Avenue Doutor Tito s/n, Jardim Panorama, 86400-000 Jacarezinho, PR, Brazil