1. Research Center for Aquatic Breeding, National Research Institute of Aquaculture, Fisheries Research Agency, 224 Hiruda, Tamaki-cho, Watarai, Mie 519-0423, Japan
2. Marine Biological Science, Faculty of Bio-resources, Mie University Graduate School, 1577 Kurimamachiya-cho, Tsu City, Mie 514-8507, Japan
3. Marine Farm Laboratory Limited Company, 309 Takahiro Tachibaura Otsuki-cho, Hata-gun, Kochi 788-0352, Japan