1. Achucarro Basque Center for Neuroscience, CIBERNED and Departamento de Neurociencias de la Universidad del País Vasco UPV/EHU, E-48940 Leioa, Spain
2. IKERBASQUE, Basque Foundation for Science, 48011 Bilbao, Spain
3. Weill Cornell Medical College, Cornell University, New York, NY 10065, USA