1. Institute of Biomedical and Biomolecular Science, University of Portsmouth, PO1 2DY Portsmouth, UK
2. Institut de Génétique et Développement,
CNRS, UMR6290, Faculté de Médecine, Université de Rennes 1, 35000 Rennes, France
3. Institute of Marine Sciences, School of Biological Sciences, University of Portsmouth, P04 9LY Portsmouth, UK