1. Laboratory of Marine Organism Taxonomy and Phylogeny, Qingdao Key Laboratory of Marine Biodiversity and Conservation, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
2. Ocean College of Tangshan Normal University, Tangshan 063000, China
3. Guangxi Key Laboratory of Beibu Gulf Marine Biodiversity Conservation, College of Marine Sciences, Beibu Gulf University, Qinzhou 535011, China
4. Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519082, China
5. University of Chinese Academy of Sciences, Beijing 100049, China