1. College of Life Sciences, Northwest A&F University, Yangling, Shaanxi 712100, China
2. Department of Plant Pathology, Washington State University, Pullman, WA 99164-6430, USA
3. USDA-ARS, Wheat Genetics Quality, Physiology, and Disease Research Unit, Pullman, WA 99164-6430, USA
4. College of Plant Protection, Northwest A&F University, Yangling, Shaanxi 712100, China