1. Department of Animal Science, University of Connecticut, Storrs, CT 06269, USA
2. Center of Excellence for Vaccine Research, University of Connecticut, Storrs, CT 06269-2191, USA
3. Department of Biomedical Sciences, Division of Infectious Diseases, Tufts Cummings School of Veterinary Medicine, North Grafton, MA 01536, USA
4. Department of Allied Health Sciences, University of Connecticut, Storrs, CT 06269, USA
5. Plum Island Animal Disease Center, USDA-APHIS/VS/FADDL, Greenport, NY 11944, USA