1. Division of Biotechnology, Ferdowsi University of Mashhad, Mashhad, Iran
2. Biotechnology Research Center, Pharmaceutical Technology Institute, Mashhad University of Medical Sciences, Mashhad, Iran
3. Department of Biotechnology, School of Pharmacy, Mashhad University of Medical Sciences, Mashhad, Iran
4. Department of basic sciences, Faculty of Veterinary Medicine, Ferdowsi University of Mashhad, Mashhad, Iran
5. Division of industrial and environmental biotechnology, National Institute of Genetic Engineering and Biotechnology, Tehran, Iran