1. a Debre Zeit Agricultural Research Center , P.O. Box 32, Debre Zeit, Ethiopia
2. b International Institute of Tropical Agriculture (IITA)—Malawi , Chitedze Research Station, P.O. Box 30258, Lilongwe, Malawi
3. c Department of Plant Breeding and Genetics , 240 Emerson Hall, Ithaca, NY, 14853, USA
4. d Cornell University , Ithaca, NY, 14853, USA