1. Department of Biodiversity and Conservation Biology, Swiss Federal Research Institute WSL, Zürcherstrasse 111, CH-8903 Birmensdorf, Switzerland
2. Department of Plant, Soil and Entomological Sciences, University of Idaho, 875 Perimeter Drive MS 2339, Moscow, ID 83844
3. Agroscope Institute for Sustainability Sciences ISS, Reckenholzstrasse 191, CH-8046 Zurich, Switzerland
4. Department of Ecology and Evolutionary Biology, University of Connecticut, Unit 3043, 75 North Eagleville Road, Storrs, CT 06269