1. Irrigated Agriculture Research and Extension Center, Washington State University, 24106 N. Bunn Rd., Prosser, WA 99350
2. Current address: Department of Biology, Yakima Valley Community College, 500 West Main St., Grandview, WA 98930
3. USDA-ARS, 5230 Konnowac Pass Rd., Wapato, WA 98951