1. Key Laboratory of Adaptation and Evolution of Plateau Biota, Chinese Academy of Sciences, Xining 810008, People's Republic of China (e-mail: )
2. University of the Chinese Academy of Sciences, Beijing 100049, People's Republic of China (e-mail: )
3. Department of Biology, The University of New Mexico, Albuquerque, New Mexico 87131, U.S.A (e-mail: )
4. Qinghai Key Laboratory of Animal Ecological Genomics, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining 810008, People's Republic of China (e-mail: )