1. Department of Integrative Biology, Natural Science Building, Michigan State University, East Lansing, Michigan 48824, U.S.A. (e-mail: muzzall@msu.edu) and
2. Michigan Department of Natural Resources, Lake St. Clair Fisheries Research Station, 33135 South River Road, Harrison Township, Michigan 48045, U.S.A. (email: hessenauerj1@michigan.gov)