1. Brain Science Institute; Tamagawa University; Tokyo Japan
2. JST CREST; Tokyo Japan
3. Division of Visual Information Processing; National Institute for Physiological Sciences; National Institutes of Natural Sciences; Okazaki Japan
4. Department of Physiological Sciences; SOKENDAI (The Graduate University for Advanced Studies); Okazaki Japan
5. Laboratory of Neural Circuitry; Graduate School of Brain Science; Doshisha University; Kyoto Japan
6. Present address: Faculty of Human Life Studies; Department of Food and Nutrition; Hagoromo University of International Studies; Osaka Japan