1. Department of Biomedical Science University of Sheffield Sheffield S10 2TN UK
2. Neuroscience Institute University of Sheffield Sheffield S10 2TN UK
3. School of Life Sciences University of Sussex Falmer Brighton BN1 9QG UK
4. School of Life Sciences Keele University Keele ST5 5BG UK
5. Mary Lyon Centre MRC Harwell Institute Oxfordshire UK
6. Mammalian Genetics Unit MRC Harwell Institute Oxfordshire UK