1. Wellcome Trust/Cancer Research UK - Gurdon Institute; Henry Wellcome Building of Cancer and Developmental Biology; University of Cambridge; Tennis Court Road Cambridge CB2 1QN UK
2. Department of Physiology; Development and Neuroscience; University of Cambridge; Cambridge CB2 3DY UK
3. Wellcome Trust - Medical Research Council Stem Cell Institute; University of Cambridge; Gleeson Building, Tennis Court Road Cambridge CB2 1QR UK