1. Institute for Cell & Molecular Biosciences, The Medical School; Newcastle University; Framlington Place Newcastle upon Tyne NE2 4HH UK
2. Department of Physiology, McIntyre Medical Sciences Building; McGill University; 3655 Promenade Sir William Osler Montreal Quebec Canada H3G 1Y6
3. Institute for Cellular Medicine, The Medical School; Newcastle University; Framlington Place Newcastle upon Tyne NE2 4HH UK
4. Marsico Lung Institute; University of North Carolina; Chapel Hill NC 27599 USA
5. School of Biological and Biomedical Sciences; Durham University; South Road Durham DH1 3LE UK