1. Department of Computer Science University of Bristol Bristol BS8 1UB UK
2. Department of Engineering Mathematics University of Bristol Bristol BS8 1UB UK
3. Medical Research Council Anatomical Neuropharmacology Unit Department of Pharmacology University of Oxford Oxford OX1 3TH UK
4. Oxford Parkinson's Disease Centre University of Oxford Oxford OX1 3QX UK
5. Nuffield Department of Clinical Neurosciences University of Oxford Oxford OX3 9DU UK