1. Department of Pharmacology II, Graduate School of Medicine, Osaka University, 2–2 Yamada‐oka, Suita, Osaka 565–0871, Japan
2. Department of Veterinary Pharmacology, Graduate School of Agriculture and Life Science, Osaka Prefecture University, 1–1 Gakuenchou, Sakai, Osaka 599–8531, Japan
3. Department of Pharmacology, Sapporo Medical University School of Medicine, South 1, West 17, Chuo‐ku, Sapporo 060–8556, Japan
4. Department of Otolaryngology, Graduate School of Medicine, Osaka University, 2–2 Yamada‐oka, Suita, Osaka 565–0871, Japan
5. Faculty of Pharmaceutical Sciences, Toyama Medical and Pharmaceutical University, 2630 Sugitani, Toyama 930–0194, Japan