1. Department of Biological and Medical Sciences; Oxford Brookes University; Oxford OX3 0BP UK
2. Department of Physiology, Development and Neuroscience; University of Cambridge; Cambridge CB2 3EG UK
3. School of Animal and Comparative Biomedical Sciences; University of Arizona; Tucson AZ 85721 USA
4. School of Animal Biology; University of Western Australia; 6009 Crawley Australia