1. Department of NeurophysiologyNational Institute of NeuroscienceNational Center of Neurology and Psychiatry Kodaira Tokyo Japan
2. Research Fellow of the Japan Society for the Promotion of Science Chiyoda‐ku Tokyo Japan
3. Systems Neuroscience SectionDepartment of NeurosciencePrimate Research InstituteKyoto University Inuyama Aichi Japan
4. PRESTOJapan Science and Technology Agency Kawaguchi Saitama Japan